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"Omne* res create 9unt divinse sapientise et potentise testes, diritbe felicitatis humanae : ex karum usu bonitas Creatoris ; ex pulckritudine sapientia Domini ; ex ceconomia in conservatione, proportione, renovatione, potentia majestatis elucet. Earum itaque indagatio ab hominibus sibi relictis semper aestimata ; a vere eruditis et sapientibus semper exculta ; male doctis et barbaris semper inimica fuit.” Linn.®us.

Quel que soit le principe de la vie animale, il ne faut qu’ouvrir les yeux pour voir qu’elle est le cbef-d’eeuvre de la Toute-puissance, et le but auquel se rappor- tent toutes ses operations.” Bruckner, Theorie du Systeme Animal, Leyden,


The sylvan powers

Obey our summons ; from their deepest dells The Dryads come, and throw their garlands wild And odorous branches at our feet ; the Nymphs That press with nimble step the mountain-thyme And purple heath -flower come not empty-handed,

But scatter round ten thousand forms minute

Of velvet moss or lichen, torn from rock

Or rifted oak or cavern deep : the Naiads too

Quit their loved native stream, from whose smooth face

They crop the lily, and each sedge and rush

That drinks the rippling tide: the frozen poles,

Where peril waits the bold adventurer’s tread,

The burning sands of Borneo and Cayenne,

All, all to us unlock their secret stores And pay their cheerful tribute.

J. Taylor, Norwich, 1818.






I. On a Tooth of Ceratodus and a Dinosaurian Claw from the Lower

Jurassic of Victoria, Australia. By A. Smith Woodward, LL.D., F.R.S., of the British Museum. (Plate I.) 1

II. Notes on Irish Hydrachuida ; with Descriptions of a new

Genus and Two new Species. By J. N. Halbert. (Plate II.) . . 4

III. Preliminary Descriptions of new Species of Amphipoda from

the Discovery Antarctic Expedition, 1902-1904. By Alfred O. Walker, F.L.S., F.Z.S 13

IV. Rhynchotal Notes.— XXXVIII. By W. L. Distant 18

V. On some West- African Species of Barbus. By G. A.

Boulenger, F.R.S 32

VI. Description of a new Barbus from the Uganda Protectorate.

By G. A. Boulenger, F.R.S 36

VII. Description of a new Mormyrid Fish from South Cameroon.

'By G. A. Boulenger, F.R.S ib.

VIII. Description of a new Tree-Viper from Mount Ruwenzori.

By G. A. Boulenger, F.R.S 37

IX. Alternation of Generations, Metamorphosis, and Direct

Development. By W. Wedekind 38

X. Natural History Notes from the R.I.M.S. Ship ‘Investigator,’

Capt. T. H. Heming, R.N., commanding. Series III., No. 13. l’wo new Barnacles dredged in 1905-6. By N. Annandale, D.Sc., Indian Museum, Calcutta 44





XI. Description of a new Species of Parnassius. By F. Moore,

D.Sc., F.Z.S 47

XII. On Three remarkable new Melolonthid Coleoptera from Sumatra and Borneo in the British Museum. By Gilbert J. Arrow. 48

XIII. On the Bats of the Genera Micronycteris and Gly phony cter is.

By Knud Andersen 50

XIV. Descriptions of Five new Freshwater Fishes from Sarawak,

Borneo, collected by Dr. 0. Hose. By C. Tate Regan, B.A 66

XV. Descriptions and Records of Bees. XII. By T. D. A.

Cockerell, University of Colorado 69

XVI. Descriptions of Two new Species of Acrceidce from Entebbe,

Uganda. By Emily Mary Sharpe 75


XVII. Descriptions of some new Species of Heterocera from

Tropical South America. By Herbert Druce, F.L.S, &c 77

XVIII. Notes on the Genus Hcematopota of the Family Tabanidce in tbe British Museum Collection. By Gertrude Ricardo. (Plates III.-VI.) 94

XIX. On Lamellicorn Coleoptera from Portuguese West Africa,

with Descriptions of new Species. By Gilbert J . Arrow 127

XX. Descriptions of new Mammals from Mount Ruwenzori. By

Oldfield Thomas 136

XXI. On a second Species of the Silurid Genus Mochocus. By

G. A. Boulenger, F.R.S 147

XXII. On a new Pigmy Antelope obtained by Col. J. J. Harrison

in the Semliki Forest. By Oldfield Thomas 148

XXIII. Preliminary Descriptions of new Species of Amphipoda from the Discovery Antarctic Expedition, 1902-1904. By Alfred 0. Walker, F.L.S., F.Z.S 150

XXIV. Description of a new Cyprinodont Fish of the Genus Jenynsia from Argentina. By C. Tate Regan, B.A 154

Neiv Books : The Fauna of British India, including Ceylon and Burma. Published under the authority of the Secretary of State for India in Council. Edited by Lt.-Col. C. T. Bingham, lihyn- chota. Vol. III. (Heteroptera Homoptera). By W. L. Distant.

A Synonymic Catalogue of Homoptera. Part I. Gkadid.ce.

By W. L. Distant 155

Locusts in Hungary, by W. F. Kirby






XXV. Natural History Note3 from R.I.M.S. Investigator.’

Series III., No. 10. On Mollusca from the Bay of Bengal and the Arabian Sea. By Edgar A. Smith, I.S.0 157

XXVI. Notes on the Genus Tamar rha, Wkr. [Lep. Tineina].

By the Rt. Hon. Lord Walsingham, M.A., LL.D., F.R.S 175

XXVII. Description of a new Tineid Moth infesting Cotton-pods in Egypt. By the Rt. Hon. Lord Walsingham, M.A., LL.D., F.R.S 178

XXVIII. On new Species of Kisteridce and Notices of others.

By G. Lewis, F.L.S. 180

XXIX. Rhynchotal Notes.— XXXIX. By W. L. Distant .... 191

XXX. Description of a new Species of Mangabey ( Cercocebus

Hamlyni). By R. I. Pocock, F.L.S., F.Z.S., Superintendent of the Zoological Society’s Gardens. (Plate VII.) 208

XXXI. On a new Species of Coral-infesting Crab taken by the

R.I.M.S. 1 Investigator at the Andaman Islands. By J. R. Hen- derson, M.B., F.L.S., Professor of Biology, Madras Christian College. (Plate VIII.) 211

XXXII. Three new Palaearctic Mammals. By Oldfield Thomas. 220

XXXIII. Two new Genera of small Mammals discovered by Mrs. Holms-Tarn in British East Africa. By Oldfield Thomas . . 222

XXXIV. The Morphology of the Madreporaiia. VIII. The Primary Septa of the Rugosa. By J. E. Duerden, Ph.I)., A.R.C.S. (Lond.j, Professor of Zoology, Rhodes University College, Grahams- town, Cqpe Colony 226

XXXV. Notes on the Habits of Tsetse-flies. By Dr. F. Creighton Wellman, Benguella, West Africa 242

Proceedings of the Geological Society 244


XXXVI. Natural History Notes from R.I M.S. Investigator.’ Series 111., No. 10. On Mollusca from the Bay of Bengal and the Arabian Sea. By Edgar A. Smith, I.S.0 245

XXXVII. Notes on the Genus Otomys. By R. C. Wroughton. 264

XXXVIII. On the Genus Cercocebus , with a Key to the known Species. By R. 1. Pocock, F.L.S., F.Z.S., Superintendent of the Zoological Society’s Gardens 278



XXXIX. On some Ethiopian Rhynchota, and Synonymical Notes.

By W. L. Distant 286

XL. On some African Bats and Rodents. By Oldfield Thomas. 294

XLI. New Mammals collected in North-east Africa by Mr. Zaphiro, and presented to the British Museum by W. N. McMillan, Esq.

By Oldfield Thomas, F.R.S 300

XLII. Natural History Notes from the R.I.M.S. Ship ‘Investi- gator/ Capt. T. IT. Heming, R.N. (retired), commanding. Series III.,

No. 14. Notes on the Skull of the Genus Aulastomatomorpha , with Descriptions of some new Deep-sea Fish. By R. E. Lloyd, M.B., B.Sc., Capt. I.M.S., Surgeon-Naturalist, Marine Survey of India . . 306

XLIII. The Relations of Palaeontology to Biology. By A. Smith Woodward, LL.D., F.R.S 312

New Books: A Descriptive Catalogue of the Tertiary Yertebrata of the Fayum, Egypt. By Charles William Andrews, D.Sc.

Die Tierischen Gifte. Von Edwin Stanton Faust . . 318, 320


XLIV. Brachiopod Nomenclature. By S. S. Buckman, F.G.S. . 321

XLV. The Flying-fish Problem. By Lieut.-Colonel C. D. Durnford 327

XLVI. On a new Race of Sciurus lokriodes from Burma. By J. Lewis Bonhote, M.A 338

XLVII. Descriptions of African Lepidoptera. By George T. Bethune-Baker, F.L.S., F.Z.S 339

XLVIII. Description of a new Chameleon of the Genus Rhampho- hon from Mashonaland. By G, A. Boulenger, F.R.S 346

XLIX. Description of a new Silurid Fish of the Genus Doumea, Salvage, from Angola. By G. A. Boulenger, F.R.S 317

L. On the Presence of Two Species of Anabas in the White Nile and the Bahr-el-Gebel. By G. A. Boulenger, F.R.S 348

LI. Rhynchotal Notes. XL. By W. L. Distant 349

LIE The Primary Septal Plan of the Rugosa. By R. G. Carruthers. (Plate IX.) 356

LIII. Oriental Reduviidce. By W. L. Distant 353

LIV. Note on the Type Specimen of the Bat Micronycteris microtis , Miller. By Marcus W. Lyon, Jun 371

LV. Descriptions of new Pymlidce of the Subfamilies Hydro - campince and Scopariance. By Sir George F. Hampson, Bart., B.A., F.Z.S., &c 373




LVI. Note on Doliichthys stellatus, Sauvage. By L. S. Berg (St. Petersburg) 393

LYII. Description of a new Species of Leucogobio from Korea.

By L. S. Berg (St. Petersburg) 394

Proceedings of the Geological Society 395, 39G


LVIII. On new Species of Histeridce and Notices of others. By G. Lewis, F.L.S 397

LIX. New and little-known Species of Eastern and Australian Ileterocera. By Colonel C. Swinhoe, M.A., F.L.S., &c 403

LX. On Myriolepis hibemica, a Palseoniscid Fish from the Irish Coal-Measures. By A. Smith Woodward, LL.D., F.R.S. (Plate X.) 416

LXI. Brief Diagnoses of a new Genus and Ten new Forms of Stenodermatous Bats. By Knud Andersen 419

LXII. On a new Species of Lyconus from the North-east Atlantic.

By E. W. L. Holt and L, W. Byrne 423

LXIII. Natural History Notes from the R.I.M.S. Ship ‘Investi- gator,’ Capt. T. H. Heming, R.N., commanding. Series III., No. 15. Second Preliminary Report on the Deep-sea Alcyonaria collected in the Indian Ocean. By Prof. J. Arthur Thomson, M.A., and W. D. Henderson, M.A., B.Sc., Carnegie Research Fellow, University of Aberdeen 427

LXIV. On the Land Molluscan Subgenus Ccelorus, Pilsbry. By G. K. Gude, F.Z.S , 433

LXV. Descriptions of some new Sharks in the British Museum Collection. By C. Tate Regan, B.A 435

% LXVI. Description of a new Lizard and a new Snake from Australia. By G. A. Boulenger, F.R.S 440

LXVH. Description of a new Snake of the Genus Glaucoma , from Somaliland. By G. A. Boulenger, F.R.S 441

LXVIII. Notes on South-American Rodents. By Oldfield Thomas 442

LXIX. A new Species of Ptcridium (Scopoli) from the North- east Atlantic. By L. W. Byrne . 448

LXX. A Collection of Fishes from the King River, Western Australia. By C. Tate Regan, B.A 450

LXXI. Description of a Second new Species of Mangabey ( Cerco - cebus Jumrachi). By R. I. Pocock, F.L.S., F.Z.S. , Superintendent of the Zoological Society’s Gardens. (Plate XI.) 454




LXXII. Descriptions of new Pyralidce of the Subfamilies Hydro- campirue and Scopariance. By Sir George F. Hampso.v, Bart., B.A., F.Z.S., &c 4-55

LXXIII. On a new Chameleon from Mount Ruwenzori. By G. A. Boulenger, F.R.S 473

A common British Starfish, by F. Jeffrey Bell ; A Correction, by G. T. Bethune-Baker ; Trichoniscus pygmceus , G. 0. Sars, a Woodlouse new to the British Fauna, by Richard S. Bagnall, F.E.S 473,474

Index 475

VII. Hamlyn’s Mangabey.

VIII. New species of coral-infesting crab.

IX. Early septa in Rugose corals.

X. Myriolepis hibernica.

XI. Jamrach's- Mangabey.


Plate I. Teeth of Ceratodus and Dinosaurian claws. II. Irish Hydrachnida.





per litora spargite museum.

Naiades, et eircum vitreos considite fontes :

Pollice virgineo teneros hie carpite flores :

Floribus et pictum, divae, replete canistrum.

At vos, o Nymphae Craterides, ite sub undas ;

Ite, recurvato variata corallia trunco Vellite muscosis e rupibus, et mihi conchas Ferte, Deae pelagi, et pingui conchylia succo."

N.Parthenii Giannettasi, Eel. 1.

No. 103. JULY 1906.

I. On a Tooth o/Ceratodus and a Dinosaurian Claw from the Lower Jurassic of Victoria , Australia , By A. Smith Woodward, LL.D., F.R.S., of the British Museum.

[Plate I.]

The Jurassic Vertebrate fauna of the Australian region is still almost unknown, some Ganoid fishes * and, perhaps, a few small Dinosaurian bones t being the only fossils repre- senting it hitherto described. A tooth of Ceratodus and a Dinosaurian claw discovered by Mr. W. H. Ferguson in the Lower Jurassic cliffs of Cape Patterson on the south coast of Victoria are thus of special interest. I am indebted to Prof. J. W. Gregory, F.R.S., for the opportunity of studying these specimens.

* A. S. Woodward, The Fossil Fishes of the Talbragar Beds,” Mem. Geol. Surv. N. S. Wales, Palaeont. no. 9 (1895) ; T. S. Hall, A new Genus and a new Species of Fish from the Mesozoic Pocks of Victoria,” Proc. Poy. Soc. Viet. n. 8. vol. xii. (1900) art. xvi.

t II. G. Seeley, “On Agrosaurits Macgillivrayi (Seeley), a Saurischian Reptile from the N.E. Coast of Australia,” Quart. Journ. Geol. Soc. vol. xlvii. (1891) pp. 164-165, with figs.

Ann , & Mag, N, Hist, Ser. 7. Vol, xviii. 1


Dr. A. S. Woodward on a Tooth 0/ Ceratodus

The tooth of Oeratodus (PI. I. fig. 1) is firmly fixed to a recognizable piece of the splenial bone, and is therefore proved to belong to the left side of the lower jaw. It unfor- tunately lacks the foremost denticle, but clearly agrees with the majority of the Mesozoic teeth of Ceratodus in possessing only four denticles altogether. It is thick and robust, with the grinding-surface slightly convex, but wavy, and marked by a very prominent coarse network of ridges (fig. 1). It is specially remarkable for the long and narrow shape of its crown, which is bounded on the inner side by a nearly straight margin, not angulated opposite the second or third denticle. So far as can be determined from a fragment, the foremost denticle of the tooth appears to have been relatively large, while the others rapidly decrease in size backwards. The second and third denticles are sharply compressed to an acute outer edge, and are separated by deep notches at the outer margin (fig. 1 a ), though not continued as conspicuous ridges on the crown. Their long axes are not oblique, but directed nearly at right angles to the inner margin. The fourth or hindmost denticle is comparatively blunt. Fine horizontal lines of growth are seen on the flattened inner (fig. 1 h) and outer faces of the tooth.

The specimen thu3 described differs from all the known Mesozoic teeth of Ceratodus in its narrowness, combined with the straightness of its inner margin and the direction of its second and third denticles. In these respects, it is interesting to observe, the tooth more nearly approaches that of the existing Ceratodus or JS eoceratodus of Queensland (fig. 2), and its only striking difference from the latter consists in its having four denticles instead of six. The multiplication of the denticles has already been observed in the teeth of certain sharks as they are traced onwards in time * ; the same phenomenon obviously occurs in Ceratodus .

There is, therefore, no doubt that the tooth from Cape Patterson represents a new species, which may be named Ceratodus avus. The fossil proves for the first time that the remarkable Dipnoan genus to which it belongs had already reached the Australian region so long ago as the early part of the Jurassic period. At that epoch Ceratodus was still living both in Europe f and in North America |, while it survived

* A. S. Woodward, On the Palaeontology of the Selachian Genus Notidanus, Cuvier,” Geol. Mag. [3] vol. iii. (1886) p. 257.

t Ceratodus Phillipsi , Agassiz, 1 Pech. Poiss. Foss.’ vol. iii. (1838) p. 135, pi. xix. fig. 17 j A. S. Woodward, Proc. Geol. Assoc, vol. xi. (1890) p. 292, pi. iii. fig. 5.

X Ceratodus Guenther i, O. C. Marsh, Amer. Journ. Sci. [3] vol. xv. (1878) p. 76, woodc.

Aiui,& Mc)vg.NaJb.Kist. S. 7. Vol XVIII. PI. I.

eJ. Careen. ci«l, litVi et. imp.


and a Dinosaurian Claw. 3

in the African and South American regions at least until the Cretaceous period *.

In the same rock as that from which the tooth of Ceratodus was obtained at Cape Patterson Mr. Ferguson found the terminal phalangeal bone shown in fig. 3. Among Jurassic fossils this specimen can only be compared with the claw of a carnivorous Dinosaur, and there is little doubt that it represents a genus more or less related to Megalosaurus f . The bone has decayed somewhat in the upper part of its proximal end, but is otherwise well preserved and displays its principal characters. The phalangeal is laterally com- pressed, so that its greatest transverse diameter is somewhat less than its original depth at the proximal end. The distal tapering half of the bone is only gently curved downwards, but at the same time bends slightly to the left side. The distal half of the lateral face is marked with the usual deep longitudinal groove connected with the fixing and nourishment of the horny claw which originally ensheathed the bone. The proximal end (fig. 3 a) is divided, as usual, by a median vertical ridge into two facettes, which are nearly flat. For comparison with this specimen one of the finest known Megalosaurian claws from the English Wealden is shown in fig. 4. The latter is shorter and stouter than the former, and its deep lateral groove extends further backwards ; but the general resemblance between the two fossils is very striking.

It is to be hoped that further diligent search may be made at Cape Patterson to recover the Vertebrate fauna indicated by these fragmentary fossils. The discovery of the terrestrial and freshwater life of the Australian region during the Jurassic period would supply a most important deficiency in palaeontological knowledge.


Fi(j. 1. Ceratodus avus, sp. n. ; left splenial with lower tooth, from the upper, outer (a), and inner ( b ) aspects. Lower Jurassic ; Cape Patterson, Victoria, Australia, spl., splenial bone.

Fig. 2. Ceratodus Forsteri, Krefft ; left lower tooth from the upper and outer (a) aspects. Recent; Queensland.

Fig. 3. Ungual phalange of carnivorous Dinosaur ; lateral and end (a) views. Lower Jurassic; Cape Patterson, Victoria.

Fig. 4. Ungual phalange of a Megalosaurian ; lateral and end (a) views, two thirds nat. size. Wealden ; Sussex. [Brit. Mus. no. R. 3170.]

Figs. 1-3 are of the natural size.

* Ceratodus africanus , E. Ilaug, Comptes Rendus,’ vol. cxxxviii. (1904) p. 1529 ; from Djoua, Timassanine, Sahara. Ceratodus Iheringi , F. Ameghino, Public. Univ. La Plata, no. 2 (1904), p. 10, fig. 1 ; from Patagonia.

t R. Owen, Fossil Reptilia of the Wealden and Purbeck Formations,” pt. iii. (Mon. Palseont. Soc. 1855 [1857]), p. 19, pi. x.



Mr. J. N. Halbert on Irish Hydrachnida.

II. Notes on Irish Hydrachnida ; with Descriptions of a new Genus and Two new Species. By J. N. Halbert.

[Plate II.]

The following paper contains records of some species of new or rare Hydrachnida selected from a large amount of material found in various localities in Ireland during the last five years. Of these species two appear not to have been previously described ; one of them proves to be the type of a new genus, while of the remaining species eight are here recorded for the first time from the Britannic * area.

It was originally intended to reserve the new mites for description in a general list of the Irish Hydrachnid fauna which is being prepared. Before such a list can be com- pleted, however, it is necessary to carry out some further collecting in certain parts of the country, and it seems more satisfactory to record the new species without further delay.

The most interesting of the new mites is one of which I was fortunate enough to find fully developed specimens when collecting last May in the south-west of Ireland. The species in question seems to be an extremely isolated form, possessing a combination of characters which at once distinguish it from any of the known genera. It would be easy to briefly define a new genus for the reception of this mite bv referring to the structure of a very few^ organs, such as the palps, legs, &c. ; in a group like the Hydrachnida, however, where there is such a great variety of structural detail, it seems especially necessary to rely on a combination of various characters in the formation of new genera.

I have to acknowledge the assistance of the Irish Fauna and Flora Committee supported by the Royal Society, and also of the same Committee when acting under the auspices of the Royal Irish Academy : several grants enabled me to collect in distant parts of the country.

The nomenclature used in the following list is that of the ' Tierreich ; (“ Hydrachnida? und Halacaridfie,’' Piersig and Lohmann, Lieferung 13, 1901).

* The use of the word Britannic instead of British for faunistic purposes has been proposed by Professor G. H. Carpenter, as the latter term is now so frequently used to distinguish records referring to Great Britain alone (‘ Irish Naturalist/ vol. xv. p. 13).

Mr. J. N. Halbert on Irish Hydrachnida .


MOMONIA*, gen. nov.

Diagnosis of Genus. An Hydrachnid of the family Hygro- batidse (Kramer, Wolcott, &c.), with a highly chitinized integument modified into chitinous plates. In shape re- sembling the genus Midea , with a convex dorsal area separated from a larger ventral field by a groove in which are a number of paired gland- openings. Epimeral groups close together, occupying most of the underside of the body. Genital area situated between the fourth epimera, flanked on each side by a triangular plate, in which are imbedded three genital suckers of the Hygrohates type. Palps with the penultimate segment angularly swollen on the ventral surface and armed with two stout chitinous teeth ; fifth segment ending in an exceedingly fine point. First pair of legs modified, the terminal segment deeply excavated on its upper margin, with a broad-shanked bifid claw articulating deeply in the segment, and, in the type species, with a terminal membrane. The three posterior pairs of legs are provided with swimming-hairs.

It will be seen from this short diagnosis that the genus Momonia possesses a very anomalous combination of characters which renders the placing of it in a satisfactory position in the Hydrachnid series a matter of some difficulty. On the whole, however, it shows affinities with the genera Midea , Mideopsis , &c., and it seems to me that it should be placed in an intermediate position between these genera and the Hygrohates group.

Momonia falcipalpis f, sp. n.

(PI. II. figs. 1-4.)

Male. Body slightly longer than broad, evenly rounded posteriorly, and narrowed towards the front margin, where there are two hair-papilla3. Seen from the side the dorsal outline is moderately convex and the ventral surface flat over the epimeral area ; thickness of the body dorso-ventrally about three fifths of the total length. Integument highly chitinized, with a dorsal groove running round close to the body-margin ; in this groove are placed at least six pairs of

* Momonia, or Mumonia, the ancient Latin name of the province of Munster. This name has also been used by Mr. R. Lloyd Praeger, M.R.I.A., to indicate the group of plants with a southern range in Ireland (see Proceed. Royal Irish Academy,’ vol. xxiv. 1902-1004).

t The specific name is suggested by the shape of the terminal palp- segment.

6 Mr. J. N. Halbert on Irish Hydrachnida .

eliitinous hair-bearing glands. The greater part of the dorsal area is covered by a large shield with sinuous side-margins bounded by the dorsal groove, the rounded posterior margin reaching to the end of the body. This plate is wrinkled longitudinally, and under a high magnification it is seen to have a finely shagreened appearance as well as polygonal reticulations. In front of this large shield lies a short broad plate, emarginate anteriorly, and rather less than half the breadth of the dorsal shield. The dark-pigmented eyes, separated by an interval of about 176 p, are situated close to the front margin of the body. On the inner side of each eye- group stands a conspicuous hair-papilla.

The greater part of the ventral side is occupied by the epimeral plates ; the first and second epimera are of the usual shape except that the first epimeron is very narrow and tapers inwardly into a rather fine point separated by an extremely narrow interval from the third epimera. The last are quadrilateral in outline, with the front and hinder margins sloping downwards. The fourth epimeron is extremely large and characteristic ; the inner margin is continued for a short distance in a line with that of the third epimeron, it then bends suddenly outwards and downwards in a sinuous line to near the hinder margin of the body, fusing with the chitinous integument of the sides of the body. Near the middle of the fourth epimeral area on each side is a group of long hairs. The epimera are all finely shagreened and reticulated in the same way as the dorsal shield.

The genital area lies in the anterior space between the fourth epimera ; it is flanked on each side by a long triangular plate, which carries three genital suckers, placed one behind the other, similar in structure to those found in Hygrohates and allied genera. A transverse chitinous plate, in which is imbedded the anal opening, occupies the remaining inter- epimeral space.

The capitulum is rather small, projecting downwards for part of its length beyond the ventral outline of the body ; maxillary shield measuring about 110 p in length (not in- cluding subcutaneous process) and 77 p in breadth.

The palps are small, the five segments measured along their dorsal margins are 30 p, 75 p} 50 //,, 85 p} 55 p respec- tively; segments 1, 2, and 3 may be compared with those of Mideopsxs ; 2 and 3 are furnished with a number of rather stout hairs on the dorsal surface ; 4 is the longest palp- segment, it is slightly convex dorsally, with two long fine hairs, ventral surface produced beyond the middle into a well- marked angular prominence, on which are two short stout


Mr. J. N. Halbert on Irish Hydrachnida.

teeth placed close to the inner side of the segment ; 5 is nearly as broad at base as the distal margin of 4, tapering gradually into a long sharp point; the upper and lower surfaces are each armed with a long hair and a sharp spine ; on the outer side close to the base is another spine.

First pair of legs (length about 814 p) modified, without swimming-cilia, slightly longer than the body, the segments gradually increasing in length from 1 to 5. Segment 1 very short, 2 and 3 straight, with a number of long bristles ; 4 curved, with the ventral distal margin notched ; 5 straight (length 260 p), broader than preceding segments, and nar- rowing towards the distal extremity, on which are seven or eight long hairs. Segment 6 (fig. 3) articulates with a conical projection on the penultimate segment, short, with convex sides, very deeply hollowed out on its upper distal margin ; a powerful claw-like structure with a broad shank and sharply bent bifid extremity articulates with the inner part of the excavation ; a few long hairs and a peculiar cone- shaped membrane project from the extremity of the segment. The last three pairs of legs do not present any remarkable characters ; they increase in length from before backwards and are provided with long swimming-cilia ; the terminal segments are armed with two recurved tridentate claws resembling those of Brachypoda.

During life the colour was a pale yellowish green, marked on the dorsal surface with reddish brown ; Malpighian area yellow, indicated anteriorly by four lobes arranged across the body.

Measurements .


Length of body 768

Breadth of body 691

Length of palp about 270

Length of leg i 814

Length of leg ii 704

Length of leg iii 792

Length of leg iv 868

Locality . Two fully developed examples of this species were found amongst a thick growth of Callitriche in Loos- caunagh Lough, about ten miles from Killarney, May 1905. From the peculiar modification of the first pair of legs there is no doubt that the specimens are males.

The type specimens are in the Dublin Natural History Museum (register no. 179, 1906).


Mr. J. N. Halbert on Irish Hydrachnida .

Arrhenurus octagonus , sp. n. (PI. II. fig. 5.)

Male . Colour during life red, with ill-defined darker markings on the back. In dorsal view the body is roughly octagonal in shape, the posterior half being somewhat similar in outline to the anterior. Front margin almost straight, about equal to half the width of the body ; all other margins very slightly emarginate. There are no conspicuous dorsal humps, but in the middle of the posterior margin there is a deep excavation, with a prominence on each side on which is a long hair. Dorsal furrow roughly circular in form, enclosing a comparatively small area (length 537 /I) of the middle of the back.

The appendage is short, measuring about a sixth of the entire length of the animal and about four fifths as broad ; in dorsal view mostly covered by the hinder part of the main body ; sides of the appendage hardly constricted at base, gradually narrowing inwards and blending with the hinder margin. Posterior dorsal margin with a wide excavation reaching from side to side ; posterior ventral margin slightly sinuate, pierced in the middle by a narrow deep indentation, which widens noticeably at its deepest part and reaches the base of the appendage. The petiolus is composed of two finely pointed pieces, which are closely approximated in the living mite, and project in the middle line very slightly beyond the margin of the appendage. There are five or six pairs of very short hairs on the end of the body.

Genital plates large, sinuate anteriorly, and gradually narrowing towards the sides of the body, which they do not overreach. Epimeral plates remarkably long and narrow, rather closely resembling those of A. sinuator, Muller.

Palps stoutly built, with prominent distal angles to the segments. The inner surface of the second segment seems to be without a hair-pad, but carries a few stout unfeathered bristles. Fourth segment with a long straight spine near the inner distal corner and a widely forked tactile hair on the apical margin.

The legs do not present any unusual characters ; they are rather stout, of moderate length, and the fourth segment of the last pair is without a spur.

Measurements .

Length of body (including appendage) .... 1*28 mm.

Breadth of body about 1*00 mm.

Breadth of appendage at base about 870 /r.

Length of palp about 430 /*.


Mr. J. N. Halbert on Irish Rydrachnida .

Locality. Found in a pond at Fenagh, Co. Carlow, by- Mr. Denis R. Pack-Beresford, M.R.I.A., during the month of August 1903.

Type specimen deposited in the Dublin Natural History Museum (register no. 180, 1906).

Arrhenurus Leuclcarti , Piersig.

Both sexes of this mite were collected in the same locality as the preceding species by Mr. Beresford. It is apparently one of our rarest Arrhenuri , these being the only Irish specimens that I have seen. Dr. George includes it in his Lincolnshire list, and Mr. Soar reports it from the Norfolk Broads.

Arrhenurus Neumani , Piersig.

This is another addition to the list of Irish Arrhenuri published a few years ago *. I found several specimens ( d and ? ) last year in Looscaunagh Lough in May, and also in Glendalough Lake, Connemara, in the following autumn. Mr. W. Williamson has taken it in Scotland (Trans. Edinb. Field-Nat. and Micros. Soc. Session 1905- 1906).

Arrhenurus Stechi , Koenike.

1894. Zur Hydrachniden-Synonymie,” Zool. Anz. xvii. p. 274, fig*. 5.

A male of this rare species occurred in a bog-pool almost filled with Sphagnum near Ross, Co. Galway, in September 1905. This is the smallest species of the genus as yet found in Ireland, my specimen measuring but 572 ya in length. The colour was pale yellow, with two black blotches showing through behind the epimera.

Localities. Up to the present time this species has been recorded from Switzerland, where it was found in a similar kind of locality (Moosseedorf-See bei Bern), Germany, and Norway (1899). Dr. George has recorded it from Lincoln- shire (‘ The Naturalist/ 1905, p. 25).

Medeopsis crassipesj Soar.

1904. u Two new British Water-Mites,” Journ. Quekett Micros. Club, p. 107, fig. 2.

Specimens of this interesting species were sent to me by Mr. W. F. de Vismes Kane, who collected them, as long ago

* * Zoologisclicr Anzeiger,’ xxvi. 1903, p. 272.


Mr. J. N. Halbert on Irish Hydrachnida.

as September 1899, in Upper Lough Erne, Co. Fermanagh. The specimens were mixed witli the commoner Mideopsis orbicularis and were so overlooked. I have since taken the species in Lough Gill, Co. Sligo.

* Sperchon brevirostris, Koenike.

1895. Neue Sper chon- Arten aus der Schweiz,” Rev. Suisse Zool. iii. p. 416, pi. xiii. figs. 1-2.

Pool by the Glenshelane River, near Cappoquin, in the county of Waterford, May 1900.

Localities. A local though widespread species in the west of Europe, having been recorded from Norway, Switzerland, Saxony, Alps (Rhatikon), and the Azores.

* Sper chon longirostris} Koenike.

1895. Neue Sperchon- Arten aus der Schweiz,” Rev. Suisse Zool. iii. p. 420, pi. xiii. figs. 3-6.

Two specimens were found in a stream at Ballysadare, Co. Sligo, in company with Panisus Michaeli. A third specimen was taken by my friend Mr. Dudley Westropp near Mullingar in April 1903.

Localities. Recorded from Germany (Erzgebirge), Switzer- land (Rhatikon), and Italy.

* Hygrobates calliger , Piersig.

1896. 11 Einige neue Hydrachniden Formen,” Zool. Anz. xix. p. 439.

Occurs on the River Nore, near Thomastown, June 1901.

Localities. Recorded from Norway, Saxony (Erzgebirge), Italy7' (Ticino), and Germany (Thuringen) .

* Laminipes bullata (Sig. Thor).

1899. Norske Hydrachnider, III.,” Arch. Naturv. Christian, xxi. p. 40, pi. xiii. figs. 129-137.

Pool by the side of Lough Leane, Killarney, June 1905.

Fortunately the single specimen taken is a male and shows the characteristic modification of the fourth pair of legs, as described and figured by Dr. Thor. This appears to be the first record of the species since the original record from Norway, and it seems to have been omitted from the volume of the ‘Tierreich* (1901) treating of the Hydrachnida.

* Species marled with an asterisk are recorded for the first time from the Britannic area.

Mr. J. N. Halbert on Irish Ilydrachnida.


*Laminipes scaurus (Koen.).

1892. Anmerkungen zuPiersigs Beitragen zur Hydrachnidenkimde,” Zool. Anzeiger, xv. p. 266, fig. 1.

Several males taken in bog-pools on lower slopes of Bragan Mountain, between the counties Monaghan and Tyrone, by Mr. W. F. de Vismes Kane in July 1900.

Localities. Norway and Germany (‘ Tierreich ').

*Tiphys mutatus (Piersig).

1893. Acercus brevipes, Zool. Anz. xvi. p. 394.

1901. Tiphys mutatus, Piersig (nom. nov.), Tierreich, p. 241.

Two specimens ( ? ) taken at Glenavy, on the shore of Lough Neagh, June 1902, The male appears to be unknown.

* Fiona stjordaliensis (Sig. Thor).

1900. Hydrachnologische Notizen, V.,” Nyt Mag. Naturvid. xxxviii. pp. 375-378, pi. xvii. figs. 21-24.

This species is allied to P. nodata , Muller, and P. contro- versiosa, Piersig, but differs sufficiently from both in the structure of the genital area, palps, and especially in the armature of the terminal segment of the third pair of legs in the male. The species was first described in 1896 by Dr. Thor, and was supposed by Dr. Piersig to be synonymous with P. controversiosa , but the more detailed description published in the above reference clearly shows the distinctions between the species.

The only Irish specimens examined were taken by Mr. W. F. de Vismes Kane in Drumreaske Lake, Co. Monaghan.

*Panisus Michaeli) Koen. (PI. II. fig. 6.)

1896. Zool. Anzeiger, xix. p. 356.

When in the west of Ireland in the spring of 1901 I found an Hydrachnid of the genus Panisus amongst water- plants in a small stream which flows into the sea at the head of Ballysadare Bay. On examination it agreed closely with the description of P. Michaeli , Koenike, except that the chitinous marginal plates of the dorsal surface numbered four on each side in my specimen, instead of five, as recorded for P. Michaeli. On sending drawings of the mite to Dr. Koenike, he was good enough to assure me that my species is identical with P. Michaeli . There are in reality only four marginal plates on each side in that species ; the statement that there


Mr. J. N. Halbert on Irish Hydrachnida .

were five was due to the outlines of the plates not being clearly visible at the time the preliminary description was made. There are sixteen chitinous plates on the dorsal surface, arranged as follows : A middle series, consisting of a large plate between the eyes ; behind this are three small circular plates, arranged on each side of the middle line; and, finally, a large terminal plate, sinuate in front, with the postero-lateral corners produced into pointed processes. The eight marginal plates are arranged in a line on each side of the body ; the most anterior of these sends forward a long- narrow prolongation on the outside of the eye. All of the dorsal plates are coarsely areolated towards their margins and more finely in the centres ; they are also very irregular in outline, differing considerably on each side of the body.

The species seems to be very local, and as I have seen no reference to figures, a drawing (fig. 6) of the dorsal surface is given ; the areolation of only the terminal plate is indicated.

Localities. Panisus Michaeli was first recorded from Switzerland, where it was discovered by Dr. A. D. Michael at Davos ; and Dr. Sig. Thor has recently recorded it from Norway. I have also seen a specimen collected by Mr. William Evans near Bolerno, Scotland, in the autumn of last year.

Thyas longirostrisj Piersig.

This very distinct species is of local occurrence in Ireland. I once found amongst Callitriche in a small pool near Ken- mare many specimens, some of which were very large,